Rowold , Perez-Benedico , Stojkovic , Garcia-Bertrand , Herrera . On the Bantu expansion. Gene. 2016 Nov 15;593(1):48-57. http://www.sciencedirect.com/science/article/pii/S0378111916305704 , report the discovery of Southeast African Bantu carriers of R1-M269
In West and Central Africa populations exhibit extremely high frequencies (61% to 95%) of R1b1-P25, or V88. In contrast, two of the three Southeast African Bantu populations genotyped in this study, display polymorphic levels of M269 in Central Mozambique 9% and Maputo 1%.
Rowold et al, speculate that M269 was introduced into Southeast Africa by Arabs or Western Europeans.
R1 originated in Africa and spread into Eurasia. I specifically stated the frequency of R1 among African populations throughout my 2011 paper.
Many Sub Saharan Africans carry R1b1b2. Wood et al (2009) found that Khoisan (2.2%) and Niger-Congo (0.4%) speakers carried the R-M269 y-chromosome. The Khoisan also carry RM343 (R1b) and M 198 (R1a1) (Naidoo et al., 2010) The Gonzalez et al article found that 10 out of 19 subjects in the study carried R1b1-P25 or M269. This is highly significant because it indicates that 53% of the R1 carriers in this study were M269, this finding is further proof of the widespread nature of this so-called Eurasian genes in Africa among populations that have not mated with Europeans.