We now know the DNA carried by the Black Gravettan and Magdalenian based on a recent paper: Paleolithic DNA from the Caucasus reveals core of West Eurasian ancestry, by Iosif Lazaridis, Anna Belfer-Cohen, Swapan Mallick, Nick Patterson, Olivia Cheronet, Nadin Rohland, Guy Bar-Oz, Ofer Bar-Yosef, Nino Jakeli, Eliso Kvavadze, David Lordkipanidze, Zinovi Matzkevich, Tengiz Meshveliani, Brendan J. Culleton, Douglas J. Kennett, Ron Pinhasi, David Reich https://www.biorxiv.org/content/10.1101/423079v1.full
Lazardis et al wrote: “ Furthermore, Dzudzuana shares more alleles with Villabruna-cluster groups than with other ESHG (Extended Data Fig. 5b), suggesting that this European affinity was specifically related to the Villabruna cluster, and indicating that the Villabruna affinity of PGNE populations from Anatolia and the Levant is not the result of a migration into the Near East from Europe. Rather, ancestry deeply related to the Villabruna cluster was present not only in Gravettian and Magdalenian-era Europeans3 but also in the populations of the Caucasus, by ~26kya. Neolithic Anatolians, while forming a clade with Dzudzuana with respect to ESHG (Extended Data Fig. 5a), share more alleles with all other PGNE …”
The authors report that these Eurasians carried mtDNA N and U6 and Y-Cromosome R1 like the Villabruna cluster. The mtDNA dates back to Cro-Magnon times when modern humans entered Europe.
The Khoisan founded civilization in Europe. They were the first Europeans. The Aurignacian civilization was founded by the Cro-Magnon people who originated in Africa. They took this culture to Western Europe across the Straits of Gibraltar. The Cro-Magnon people were probably Bushman/Khoi.
There have been numerous "Negroid skeletons" found in Europe. Marcellin Boule and Henri Vallois, in Fossil Man, provide an entire chapter on the Africans/Negroes of Europe Anta Diop also discussed the Negroes of Europe in Civilization or Barbarism, pp.25-68. Also W.E. B. DuBois, discussed these Negroes in the The World and Africa, pp.86-89. DuBois noted that "There was once a an "uninterrupted belt' of Negro culture from Central Europe to South Africa" (p.88).
Boule and Vallois, note that "To sum up, in the most ancient skeletons from the Grotte des Enfants we have a human type which is readily comparable to modern types and especially to the Negritic or Negroid type" (p.289). They continue, "Two Neolithic individuals from Chamblandes in Switzerland are Negroid not only as regards their skulls but also in the proportions of their limbs. Several Ligurian and Lombard tombs of the Metal Ages have also yielded evidences of a Negroid element.
Since the publication of Verneau's memoir, discoveries of other Negroid skeletons in Neolithic levels in Illyria and the Balkans have been announced. The prehistoric statues, dating from the Copper Age, from Sultan Selo in Bulgaria are also thought to protray Negroids.
In 1928 Rene Bailly found in one of the caverns of Moniat, near Dinant in Belgium, a human skeleton of whose age it is difficult to be certain, but seems definitely prehistoric. It is remarkable for its Negroid characters, which give it a reseblance to the skeletons from both Grimaldi and Asselar (p.291).
Boule and Vallois, note that "We know now that the ethnography of South African tribes presents many striking similarities with the ethnography of our populations of the Reindeer Age. Not to speak of their stone implements which, as we shall see later , exhibit great similarities, Peringuey has told us that in certain burials on the South African coast 'associated with the Aurignacian or Solutrean type industry...."(p.318-319). They add, that in relation to Bushman art " This almost uninterrupted series leads us to regard the African continent as a centre of important migrations which at certain times may have played a great part in the stocking of Southern Europe. Finally, we must not forget that the Grimaldi Negroid skeletons sho many points of resemblance with the Bushman skeletons". They bear no less a resemblance to that of the fossil Man discovered at Asslar in mid-Sahara, whose characters led us to class him with the Hottentot-Bushman group.
The Boule and Vallois research makes it clear that the Bushman expanded across Africa on into Europe via Spain as the Grimaldi people. This makes it clear that the Bushman/Khoisan people were not isolated in South Africa. The Khoisan people carry the haplogroup N. The Hadza are Bushman they carry haplogroup N.
“Specific mtDNA sites outside HVRI were also analyzed (by amplification, cloning, and sequencing of the surrounding region) to classify more precisely the ancient sequences within the phylogenetic network of present-time mtDNAs (35, 36). Paglicci-25 has the following motifs: +7,025 AluI, 00073A, 11719G, and 12308A. Therefore, this sequence belongs to either haplogroups HV or pre-HV, two haplogroups rare in general but with a comparatively high frequencies among today's Near-Easterners (35). Paglicci-12 shows the motifs 00073G, 10873C, 10238T, and AACC between nucleotide positions 10397 and 10400, which allows the classification of this sequence into the macrohaplogroupN,containing haplogroups W, X, I, N1a, N1b, N1c, and N*. Following the definition given in ref. 36, the presence of a single mutation in 16,223 within HRVI suggests a classification of Paglicci-12 into the haplogroup N*, which is observed today in several samples from the Near East and, at lower frequencies, in the Caucasus (35). It is difficult to say whether the apparent evolutionary relationship between Paglicci-25 and Paglicci-12 and those populations is more than a coincidence. Indeed, the haplogroups to which the Cro-Magnon type sequences appear to belong are rare among modern samples, and therefore their frequencies are poorly estimated. However, genetic affinities between the first anatomically modern Europeans and current populations of the Near East make sense in the light of the likely routes of Upper Paleolithic human expansions in Europe, as documented in the archaeological record (37)”. http://www.pnas.org/cgi/content/full/100/11/6593
During the Aurignacian period, the Cro magnon skeletons belong to the N haplogroup.
The Cro Magnon skeletons carried N1a,N1b,N1c and N*. It is characterized by motifs 00073G,10873C, 10238T and A4CC between nucleotide positions 10397 and 10400. Most of the skeletons carried hg N*.
It appears that the hg N was the most frequent mtDNA carried by Western European populations for over 20,000 years. This gene as discussed earlier is found primarily today outside Western Europe. The Cro Magnon people were mainly hunter-gathers.
Haak et al. found that the twenty-four samples included haplogroups H or V, T, K, J , N1a and U3. The frequency of N1a among ancient samples ranged from 8% to 42%.
Haak et al found that the first Neolithic farmers did not have a strong genetic influence on modern European female lineages. These researchers found that the farmers were predominately HG N1a. This is interesting because Brace et al found that the craniofacial features of these early European farmers and the Natufians plotted with Sub-Saharan groups, just like the Aurignacians. The existence of the hg N in western Europe from 24,000-7500 kya show continuity between the Pleistocene and Neolithic western Eurasians who carried hg N.